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Males 2 |
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Females 12 |
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2 (1)
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Tibial apophysis narrow; body length smaller or equal 4.0 mm; prosoma length smaller or equal 2.0 mm Haplodrassus minor (O. Pickard-Cambridge, 1879)
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Tibial apophysis different and/or body length larger 3 |
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3 (2) |
Tibial apophysis distally broadened 4 |
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Tibial apophysis distally not broadened 8 |
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4 (3)
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Distal bulbal apophysis with strikingly serrated margin Haplodrassus kulczynskii Lohmander, 1942
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Distal bulbal apophysis different 5 |
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5 (4) |
Distal bulbal apophysis long and slender 6 |
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Distal bulbal apophysis broader and 9 |
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6 (5)
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Pedipalp as fig.; tibial apophysis distally hardly broadened Haplodrassus bohemicus Miller & Buchar, 1977
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Different 7 |
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7 (6)
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Distal bulbal apophysis distinctly keeled Haplodrassus signifer (C. L. Koch, 1839)
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Distal bulbal apophysis without keel Haplodrassus aenus Thaler, 1984
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8 (3)
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Plate-like bulge of distal bulbal apophysis situated distally, overtops tip of apophysis; tibial apophysis distinctly broadened distally Haplodrassus soerenseni (Strand, 1900)
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Plate-like bulge of distal bulbal apophysis situated basally; tibia apophysis only little broadened Haplodrassus umbratilis (L. Koch, 1866)
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9 (5) |
Distal bulbal apophysis distinctly broadened distally 10 |
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Distal bulbal apophysis not broadened distally and with 2 tooth-like structures Haplodrassus dalmatensis (L. Koch, 1866)
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10 (9)
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Distal bulbal apophysis ± straight, basally with a tooth; tibial apophysis as fig. Haplodrassus cognatus (Westring, 1861)
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 Palpal tibia, dorsal view (Miller & Buchar 1977)
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Distal bulbal apophysis ± rectangularly bent, without tooth 11 |
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11 (10)
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Tibial apophysis large, reaching 1/2 of bulbal length, distinctly keeled Haplodrassus silvestris (Blackwall, 1833)
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Tibial apophysis small, reaching about 1/3 of bulbal length, without distinct keel Haplodrassus moderatus (Kulczyński, 1897)
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12 (1) |
Epigyne with several transverse folds 13 |
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Different 14 |
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13 (12)
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Bar of anterior epigynal margin narrow; depressed area between the sclerotised pockets narrow Haplodrassus silvestris (Blackwall, 1833)
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Bar of anterior epigynal margin narrow as broad as epigyne; depressed area between sclerotised pockets broad and with strong indentations Haplodrassus cognatus (Westring, 1861)
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14 (12) |
Sclerotised epigynal pockets with shallow protrusion 15 |
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Sclerotised epigynal pockets without protrusion 17 |
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15 (14)
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Protrusion very elongated Haplodrassus kulczynskii Lohmander, 1942
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Different 16 |
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16 (15)
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Protrusion slantingly forward-directed Haplodrassus umbratilis (L. Koch, 1866)
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Protrusions directed more towards median region Haplodrassus soerenseni (Strand, 1900)
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17 (14) |
Margins of sclerotised, black epigynal pockets parallel or slightly roundish curved 18 |
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Margins of sclerotised, black epigynal pockets strongly converging or diverging posteriorly 21 |
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18 (17) |
Margins on the inner side roundish curved 19 |
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Margins on the inner side straight 20 |
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19 (18)
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Epigyne and vulva as figs. Haplodrassus signifer (C. L. Koch, 1839)
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Epigyne and vulva as figs. Haplodrassus bohemicus Miller & Buchar, 1977
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20 (18)
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Epigyne and vulva as figs. Haplodrassus aenus Thaler, 1984
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Epigyne and vulva as figs. Haplodrassus minor (O. Pickard-Cambridge, 1879)
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21 (17)
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Margins diverging towards posterior Haplodrassus dalmatensis (L. Koch, 1866)
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Margins converging towards posterior Haplodrassus moderatus (Kulczyński, 1897)
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| 1. Haplodrassus alexeevi Ponomarev & Shmatko, 2017 |
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| 2. Haplodrassus caspius Ponomarev & Belosludtsev, 2008 |
 Pedipalp, retrolateral view (Kovblyuk et al. 2013d)
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 Pedipalp, ventral view (Kovblyuk et al. 2013d)
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 Pedipalp, prolateral view (Kovblyuk et al. 2013d)
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 Pedipalp, retrolateral view (Chatzaki 2021)
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 Pedipalp, ventral view (Chatzaki 2021)
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 Bulbus, dorsal view (Kovblyuk et al. 2013d)
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 Bulbus, apical view (Kovblyuk et al. 2013d)
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 Tip of bulbus, dorsal view (Chatzaki 2021)
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 Epigyne, ventral view (Ponomarev et al. 2008)
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 Epigyne, ventral view (Kovblyuk et al. 2013d)
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 Eyes, frontal view (Kovblyuk et al. 2013d)
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| 3. Haplodrassus caucasius Ponomarev & Dvadnenko, 2013 |
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| 4. Haplodrassus concertor (Simon, 1878) |
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| 5. Haplodrassus crassipes (Lucas, 1846) |
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| 6. Haplodrassus creticus (Roewer, 1928) |
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| 7. Haplodrassus dentifer Bosmans & Abrous, 2018 |
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| 8. Haplodrassus ibericus Melic, Silva & Barrientos, 2016 |
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| 9. Haplodrassus invalidus (O. Pickard-Cambridge, 1872) |
 Pedipalp, ventral view (Levy 2004)
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 Pedipalp, ventral view (Kovblyuk et al. 2012b)
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 Pedipalp, prolateral view (Bosmans et al. 2018)
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 Pedipalp, retrolateral view (Bosmans et al. 2018)
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 Pedipalp, ventral view (Bosmans et al. 2018)
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 Bulbus, apical view (Kovblyuk et al. 2012b)
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 Palpal tibia, dorso-lateral view (Bosmans et al. 2018)
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 Tibial apophysis (Levy 2004)
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 Tibial apophysis, retrolateral view (Kovblyuk et al. 2012b)
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 Epigyne (Levy 2004)
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 Epigyne (Bosmans et al. 2018)
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 Epigyne (Chatzaki & Van Keer 2019)
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 Vulva, dorsal view (Levy 2004)
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 Vulva, ventral view (Bosmans et al. 2018)
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| 10. Haplodrassus ivlievi Ponomarev, 2015 |
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| 11. Haplodrassus longivulva Bosmans & Hervé, 2018 |
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| 12. Haplodrassus lyndae Abrous & Bosmans, 2018 |
 Pedipalp, prolateral view (Bosmans et al. 2018)
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 Pedipalp, retrolateral view (Bosmans et al. 2018)
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 Pedipalp, ventral view (Bosmans et al. 2018)
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 Pedipalp, ventral view (Bosmans et al. 2018)
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 Pedipalp, ventral view (Breitling 2020)
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 Embolus, dorsal view (Bosmans et al. 2018)
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 Embolic division, ventral view (Bosmans et al. 2018)
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 Palpal tibia, retrolateral view (Breitling 2020)
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 Tibial apophysis, dorso-lateral view (Bosmans et al. 2018)
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 Tibial apophysis, dorso-lateral view (Bosmans et al. 2018)
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 Epigyne (Bosmans et al. 2018)
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 Epigyne (Bosmans et al. 2018)
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 Vulva, dorsal view (Bosmans et al. 2018)
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 Vulva, ventral view (Bosmans et al. 2018)
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| 13. Haplodrassus macellinus (Thorell, 1871) |
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 Pedipalp, ventral view (Simon 1878)
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 Pedipalp, retrolateral view (Lecigne unpubl.)
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 Pedipalp, ventral view (Lecigne unpubl.)
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 Pedipalp, retrolateral view (Bosmans et al. 2018)
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 Pedipalp, ventral view (Bosmans et al. 2018)
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 Pedipalp, retrolateral view (Lecigne 2018b)
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 Pedipalp, ventral view (Lecigne 2018b)
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 Pedipalp, ventro-prolateral view (Lecigne 2018b)
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 Palpal tibia (O. Pickard-Cambridge 1874)
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 Palpal tibia, dorso-lateral view (Bosmans et al. 2018)
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 Palpal tibia, retrolateral view (Lecigne 2018b)
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 Epigyne (Di Franco 1996)
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 Epigyne, ventral view (Simon 1878)
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 Epigyne (Bosmans et al. 2018)
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 Vulva, dorsal view (Bosmans et al. 2018)
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 Vulva, ventral view (Bosmans et al. 2018)
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| 14. Haplodrassus mediterraneus Levy, 2004 |
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| 15. Haplodrassus morosus (O. Pickard-Cambridge, 1872) |
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| 16. Haplodrassus nabozhenkoi Ponomarev, 2023 |
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| 17. Haplodrassus nigroscriptus (Simon, 1909) |
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 Pedipalp, dorsal view (Bosmans et al. 2018)
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 Pedipalp, retrolateral view (Bosmans et al. 2018)
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 Pedipalp, ventral view (Bosmans et al. 2018)
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 Palpal tibia, dorso-lateral view (Bosmans et al. 2018)
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 Palpal tibia, dorso-lateral view (Bosmans et al. 2018)
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 Pedipalp, lateral view (Bosmans et al. 2018)
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 Epigyne (Bosmans et al. 2018)
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 Epigyne (Bosmans et al. 2018)
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 Vulva, ventral view (Bosmans et al. 2018)
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 Vulva, dorsal view (Bosmans et al. 2018)
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 Vulva, ventral view (Bosmans et al. 2018)
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| 18. Haplodrassus omissus (O. Pickard-Cambridge, 1872) |
 Pedipalp, ventral view (Bosmans & Van Keer 2012)
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 Pedipalp, ventral view (Bosmans & Van Keer 2012)
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 Pedipalp, prolateral view (Bosmans et al. 2018)
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 Pedipalp, retrolateral view (Bosmans et al. 2018)
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 Pedipalp, ventral view (Bosmans et al. 2018)
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 Pedipalp, ventral view (Bosmans et al. 2018)
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 Apical part of bulbus (Denis 1956a)
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 Embolic division, ventral view (Bosmans et al. 2018)
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 Palpal tibia, dorsal view (Denis 1956a)
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 Palpal tibia, dorso-lateral view (Bosmans et al. 2018)
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 Palpal tibia, dorso-lateral view (Bosmans et al. 2018)
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 Vulva, ventral view (Bosmans et al. 2018)
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 Epigyne (Chatzaki et al. 2002a)
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 Epigyne, ventral view (Denis 1956a)
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 Epigyne (Bosmans et al. 2018)
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 Epigyne (Bosmans et al. 2018)
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 Vulva (Chatzaki et al. 2002a)
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 Vulva, dorsal view (Bosmans et al. 2018)
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 Vulva, dorsal view (Bosmans et al. 2018)
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| 19. Haplodrassus orientalis (L. Koch, 1866) |
 Pedipalp, retrolateral view (Kovblyuk et al. 2012b)
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 Pedipalp, ventral view (Kovblyuk et al. 2012b)
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 Pedipalp, ventral view (Ponomarev & Tsvetkov 2006)
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 Pedipalp, retrolateral view (Ponomarev & Tsvetkov 2006)
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 Pedipalp, dorsal view (Naumova et al. 2021)
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 Pedipalp, prolateral view (Naumova et al. 2021)
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 Pedipalp, retrolateral view (Naumova et al. 2021)
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 Pedipalp, ventral view (Naumova et al. 2021)
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 Pedipalp, retrolateral view (Seyyar et al. 2022a)
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 Pedipalp, prolateral view (Seyyar et al. 2022a)
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 Pedipalp, dorso-retrolateral view (Seyyar et al. 2022a)
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 Pedipalp, ventral view (Seyyar et al. 2022a)
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 Pedipalp, ventral view (Seyyar et al. 2022a)
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 Bulbus, apical view (Kovblyuk et al. 2012b)
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 Tip of pedipalp, ventral view (Esyunin & Tuneva 2020)
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 Pedipalp (Bosmans et al. 2018)
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 Epigyne, ventral view (Kovblyuk et al. 2012b)
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 Epigyne, ventral view (Ponomarev & Tsvetkov 2006)
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 Epigyne (L. Koch 1866)
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 Epigyne (Bosmans et al. 2018)
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 Epigyne (Naumova et al. 2021)
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 Epigyne (Seropian et al. 2023b)
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 Vulva, dorsal view (Kovblyuk et al. 2012b)
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 Vulva, dorsal view (Bosmans et al. 2018)
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 Vulva, dorsal view (Bosmans et al. 2018)
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 Vulva, dorsal view (Bosmans et al. 2018)
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 Vulva, ventral view (Bosmans et al. 2018)
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 Vulva, dorsal view (Naumova et al. 2021)
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 Vulva, dorsal view (Seropian et al. 2023b)
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| 20. Haplodrassus ovatus Bosmans & Hervé, 2018 |
 Pedipalp, prolateral view (Bosmans et al. 2018)
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 Pedipalp, retrolateral view (Bosmans et al. 2018)
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 Pedipalp, ventral view (Bosmans et al. 2018)
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 Pedipalp, ventral view (Bosmans et al. 2018)
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 Embolic apophyis, ventral view (Bosmans et al. 2018)
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 Embolic apophyis, ventral view (Bosmans et al. 2018)
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 Palpal tibia, dorso-lateral view (Bosmans et al. 2018)
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 Palpal tibia, dorso-lateral view (Bosmans et al. 2018)
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 Epigyne (Bosmans et al. 2018)
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 Epigyne (Bosmans et al. 2018)
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 Vulva, dorsal view (Bosmans et al. 2018)
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 Vulva, ventral view (Bosmans et al. 2018)
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 Vulva, ventral view (Bosmans et al. 2018)
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| 21. Haplodrassus ovtchinnikovi Ponomarev, 2008 |
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| 22. Haplodrassus ponomarevi Kovblyuk & Seyyar, 2009 |
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| 23. Haplodrassus pseudosignifer Marusik, Hippa & Koponen, 1996 |
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| 24. Haplodrassus pugnans (Simon, 1880) |
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| 25. Haplodrassus rhodanicus (Simon, 1914) |
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| 26. Haplodrassus rufipes (Lucas, 1846) |
 Pedipalp, ventral view (Simon 1914)
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 Pedipalp, retrolateral view (Bosmans et al. 2018)
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 Pedipalp without embolic part, ventral view (Bosmans et al. 2018)
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 Pedipalp, ventral view (Bosmans et al. 2018)
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 Pedipalp, ventral view (Breitling 2020)
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 Embolic division, ventral view (Bosmans et al. 2018)
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 Palpal tibia, retrolateral view (Bosmans et al. 2018)
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 Palpal tibia, dorso-lateral view (Bosmans et al. 2018)
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 Palpal tibia, retrolateral view (Breitling 2020)
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 Epigyne (Simon 1914)
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 Epigyne (Bosmans et al. 2018)
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 Epigyne (Bosmans et al. 2018)
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 Epigyne, variation (Simon 1914)
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 Vulva, dorsal view (Bosmans et al. 2018)
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 Vulva (Bosmans et al. 2018)
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| 27. Haplodrassus securifer Bosmans & Abrous, 2018 |
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| 28. Haplodrassus stuxbergi (L. Koch, 1879) |
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| 29. Haplodrassus triangularis Bosmans, 2018 |
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| 30. Haplodrassus typhon (Simon, 1878) |
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| 31. Haplodrassus umbratilis gothicus Lohmander, 1942 |
 -- Caption missing -- (WSC 2024) |
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