Linyphiidae: Walckenaeria Blackwall, 1833

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1
Males

  2  

- Females

  24  

2
(1)

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Head region tower like extended

Walckenaeria acuminata Blackwall, 1833

female

- Different

  3  

3
(2)

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cp

Prosomal process narrow, long and forked; pedipalp as fig.

Walckenaeria furcillata (Menge, 1869)

female

- Different

  4  

4
(3)
Cephalic lobe distinct, ± as high as long, in some species almost as high as long

  5  

- Cephalic lobe half as high as long at most, horn-shaped or cone-shaped process or missing

  12  

5
(4)
Cephalic lobe divided into two parts by deep constriction

  6  

- Different

  8  

6
(5)

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cp

Anterior part of cephalic lobe distinctly larger than posterior part

Walckenaeria capito (Westring, 1861)

female

- Anterior part of cephalic lobe equal in size or smaller than posterior part

  7  

7
(6)

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cp

Anterior part of cephalic lobe smaller than posterior part; tibial apophysis with ventral branch tapering to a tip, dorsal branch with three blunt teeth

Walckenaeria cucullata (C. L. Koch, 1836)

female

-

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Anterior part of cephalic lobe approximately equal in size as posterior part; tibial apophysis without dorsal branch, 3-4 teeth on stem of inner apophysis; smaller than cucullata

Walckenaeria languida (Simon, 1915)

female

8
(5)

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cp

Cephalic lobe distinctly marked off; pedipalp as fig.

Walckenaeria mitrata (Menge, 1868)

female

- Different

  9  

9
(8)

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Cephalic lobe distinctly higher than long; pedipalp as fig.

Walckenaeria nodosa O. Pickard-Cambridge, 1873

female

- Different

  10  

10
(9)

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cp

Cephalic lobe as fig.; embolus spiral, approximately 1.5 loops, conducted by membranous conductor at tip; tibial apophysis pointed and bent

Walckenaeria incisa (O. Pickard-Cambridge, 1871)

female

- Cephalic lobe as fig.; embolus spiral, approximately one loop; tibial apophysis different

  11  

11
(10)

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Cephalic lobe as fig.; embolus narrowly twisted

Walckenaeria antica (Wider, 1834)

female

-

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Embolus widely twisted; otherwise similar to antica and hardly distinguishable without comparison

Walckenaeria alticeps (Denis, 1952)

female

12
(4)

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cp

Between anterior median eyes and posterior median eyes a tuft of bent plumose hairs; without cephalic lobe; pedipalp as fig.

Walckenaeria monoceros (Wider, 1834)

female

- Different

  13  

13
(12)
Prosoma with semi-high cephalic lobe

  15  

- Different

  14  

14
(13)
Prosoma with narrow, horn-shaped or cone-shaped processes between eyes

  18  

- Prosoma without conspicuous processes

  22  

15
(13)

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cp

Prosoma step-like elevated anteriorly, in the middle of dorsal line with a distinct depression; tibia with two equally long apophyses

Walckenaeria atrotibialis (O. Pickard-Cambridge, 1878)

female

- Different

  16  

16
(15)

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cp

Prosoma anterior to cephalic lobe with a horn-shaped structure consisting of 15-20 merged hairs; pedipalp as fig.;

Walckenaeria stylifrons (O. Pickard-Cambridge, 1875)

female

- Prosoma without this structure

  17  

17
(16)

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Tibial apophysis long, bent, tapering to a tip, on its base another pointed process

Walckenaeria obtusa Blackwall, 1836

female

-

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Tibial apophysis long, bent, bluntly tapering, on its base another bent, blunt process; prosoma as fig.; legs completely spineless

Walckenaeria simplex Chyzer, 1894

female

18
(14)
Between the eyes a bipartite or distally broadened cone-shaped tubercle

  19  

- Between the eyes a single cone-shaped tubercle, which bears longer hairs distally

  21  

19
(18)

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cp

Ends of cone-shaped tubercle forwardly bent; pedipalp with large, spiral embolus

Walckenaeria kochi (O. Pickard-Cambridge, 1873)

female

- Ends of cone-shaped tubercle not bent

  20  

20
(19)

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cp

Tibial apophysis with 2 crossing processes

Walckenaeria clavicornis (Emerton, 1882)

female

-

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Tibial apophysis with 2 bent processes, not crossing

Walckenaeria unicornis O. Pickard-Cambridge, 1861

female

21
(18)

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Cone-shaped tubercle protruding forwards; embolus grooved near tip

Walckenaeria cuspidata Blackwall, 1833

female

-

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Cone-shaped tubercle projecting upwards: pedipalp with non-grooved embolus and claw-shaped terminal apophysis

Walckenaeria corniculans (O. Pickard-Cambridge, 1875)

female

22
(14)

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Pedipalp with small paracymbium; tibial apophysis rounded and simple

Walckenaeria vigilax (Blackwall, 1853)

female

- Pedipalp with large paracymbium; tibial apophysis complex

  23  

23
(22)

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Paracymbium ± U-shaped; tibial apophysis distally with 2 long processes

Walckenaeria nudipalpis (Westring, 1851)

female

-

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Paracymbium ± elongated; tibial apophysis distally with one long process

Walckenaeria dysderoides (Wider, 1834)

female

24
(1)

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Epigyne with scapus; Vulva as fig.

Walckenaeria dysderoides (Wider, 1834)

- Epigyne without distinct scapus

  25  

25
(24)

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Epigyne with 2 tongue-shaped plates, reaching far beyond epigastric furrow

Walckenaeria clavicornis (Emerton, 1882)

male

- Different

  26  

26
(25)

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cp

Margins of lateral epigynal plates ± parallel, Y-like diverging anteriorly

Walckenaeria unicornis O. Pickard-Cambridge, 1861

male

- Different

  27  

27
(26)
Median epigynal plate nose-shaped, reaching beyond epigastric furrow

  28  

- Different

  29  

28
(27)

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cp

Epigyne and vulva as fig.

Walckenaeria cucullata (C. L. Koch, 1836)

male

-

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Epigyne and vulva as fig.; smaller than cucullata

Walckenaeria languida (Simon, 1915)

male

29
(27)
Median epigynal plate semicircular, reaching beyond epigastric furrow

  30  

- Different

  33  

30
(29)
Median epigynal plate conspicuously sclerotised on posterior epigynal margin

  31  

- Part of median epigynal plate which reaches beyond epigastric furrow without conspicuously margin

  32  

31
(30)

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cp

Median epigynal plate anteriorly distinctly marked off as roundish oval plate

Walckenaeria furcillata (Menge, 1869)

male

-

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cp

Median epigynal plate not marked off anteriorly

Walckenaeria incisa (O. Pickard-Cambridge, 1871)

male

32
(30)

world

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cp

Median epigynal plate broadly projecting beyond epigastric furrow

Walckenaeria stylifrons (O. Pickard-Cambridge, 1875)

male

-

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cp

Median epigynal plate roundly projecting beyond epigastric furrow

Walckenaeria nodosa O. Pickard-Cambridge, 1873

male

33
(29)
Length of marked off part of median epigynal plate approximately 3/4 of epigynal length

  34  

- Length of marked off part of median epigynal plate 1/2 of epigynal length at most

  36  

34
(33)

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Median epigynal area hourglass-shaped, widest posteriorly

Walckenaeria kochi (O. Pickard-Cambridge, 1873)

male

- Different

  35  

35
(34)

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cp

Median epigynal area widest anteriorly

Walckenaeria mitrata (Menge, 1868)

male

-

world

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cp

Epigyne and vulva as fig.

Walckenaeria simplex Chyzer, 1894

male

36
(33)

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Median epigynal plate distinctly marked off from remaining epigynal area

Walckenaeria capito (Westring, 1861)

male

- Median epigynal plate merging with remaining epigynal on anterior margin at least

  37  

37
(36)

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Eye region of prosoma strongly elevated

Walckenaeria acuminata Blackwall, 1833

male

- Different

  38  

38
(37)
Receptaculum seminis visible through epigynal plate

  39  

- Receptaculum seminis not visible through anterior or median epigynal part

  40  

39
(38)

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Median epigynal plate broad; roundish, separated entrance ducts visible through epigyne

Walckenaeria nudipalpis (Westring, 1851)

male

-

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cp

Median epigynal plate trapezoid; entrance ducts Y-shaped

Walckenaeria obtusa Blackwall, 1836

male

40
(38)
Receptaculum seminis only visible through anterior or median epigynal area

  41  

- Entrance ducts, like receptaculum seminis ± distinctly visible

  43  

41
(40)

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Anterior margin of median epigynal plate bent inwards

Walckenaeria atrotibialis (O. Pickard-Cambridge, 1878)

male

- Different

  42  

42
(41)

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Epigynal area almost twice as long as wide; receptacula seminis visible through epigyne, separated by 1.5 diameter

Walckenaeria vigilax (Blackwall, 1853)

male

-

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cp

Epigynal area ± as long as wide; receptacula seminis visible through epigyne, separated by 0.5 diameter

Walckenaeria corniculans (O. Pickard-Cambridge, 1875)

male

43
(40)
Posterior epigynal margin rounded, receptaculum seminis and entrance ducts H-shaped (shining through epigyne)

  44  

- Posterior epigynal margin straight, structures visible through epigyne not H-shaped

  45  

44
(43)

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Epigyne and vulva as fig.

Walckenaeria alticeps (Denis, 1952)

male

-

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Epigyne as fig.; very similar to alticeps

Walckenaeria antica (Wider, 1834)

male

45
(43)

world

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Median epigynal plate ± trapezoid, its anterior margin bent inwards

Walckenaeria cuspidata Blackwall, 1833

male

-

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Median epigynal plate only hinted

Walckenaeria monoceros (Wider, 1834)

male

Further taxa

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1. Walckenaeria abantensis Wunderlich, 1995

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2. Walckenaeria aksoyi Seyyar, Demir & Türkes, 2008

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3. Walckenaeria angelica Millidge, 1979

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4. Walckenaeria angustifrons (Simon, 1884)

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5. Walckenaeria baborensis Bosmans, 1993

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6. Walckenaeria bifasciculata Tanasevitch, 1987

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7. Walckenaeria brucei (Tullgren, 1955)

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8. Walckenaeria christae Wunderlich, 1995

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9. Walckenaeria cirriceps Thaler, 1996

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10. Walckenaeria claviloba Wunderlich, 1995

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11. Walckenaeria coniceps Thaler, 1996

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12. Walckenaeria cretaensis Wunderlich, 1995

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13. Walckenaeria crocata (Simon, 1884)

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14. Walckenaeria cuspidata obsoleta Chyzer & Kulczyński, 1894

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15. Walckenaeria cyprusensis Wunderlich, 1995

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16. Walckenaeria dalmasi (Simon, 1915)

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17. Walckenaeria danismani Esyunin, Vlasov & Ustinova, 2024

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18. Walckenaeria erythrina (Simon, 1884)

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19. Walckenaeria extraterrestris Bosmans, 1993

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20. Walckenaeria hamus Wunderlich, 1995

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21. Walckenaeria heimbergi Bosmans, 2007

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22. Walckenaeria inflexa (Westring, 1861)

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23. Walckenaeria insperata Millidge, 1979

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24. Walckenaeria kabyliana Bosmans, 1993

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25. Walckenaeria karpinskii (O. Pickard-Cambridge, 1873)

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26. Walckenaeria kazakhstanica Eskov, 1995

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27. Walckenaeria korobeinikovi Esyunin & Efimik, 1996

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28. Walckenaeria lepida (Kulczyński, 1885)

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29. Walckenaeria mariannae Bosmans, 1993

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30. Walckenaeria neglecta Bosmans, 1993

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31. Walckenaeria picetorum (Palmgren, 1976)

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32. Walckenaeria plumata Millidge, 1979

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33. Walckenaeria pyrenaea (Denis, 1952)

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34. Walckenaeria suspecta (Kulczyński, 1882)

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35. Walckenaeria tenuitibialis Bosmans, 1993

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36. Walckenaeria torta Bosmans, 1993

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37. Walckenaeria turbulenta Bosmans, 1993

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38. Walckenaeria vilbasteae Wunderlich, 1980

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39. Walckenaeria westringi Strand, 1903

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40. Walckenaeria wunderlichi Tanasevitch, 1983