Haplodrassus signifer (C. L. Koch, 1839)
Description
Male palp with distinctly keeled distal bulbal apophysis. Prosoma red-brown, eye region darker. Chelicerae darker than prosoma. Legs brighter than prosoma. Opisthosoma provided with copper-coloured hairs.
Body length male: 4.7-7.4 mmBody length female: 5.1-11.8 mm
Additional information
On dry meadows, on sand and stony debris, lawns, heathland, open forests and damp areas, up to 3000 m in the Alps, in litter and under stones during the day
Distribution
Phenology
| Jan | Feb | Mar | Apr | May | Jun | Jul | Aug | Sep | Oct | Nov | Dec |
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Figures
Pedipalp, ventral view
(Brændegaard 1966)
Pedipalp, ventral view
(Noordam 1992)
Pedipalp
(Noordam 1992)
Pedipalp
(Roberts 1995)
Pedipalp
(Locket & Millidge 1951)
Pedipalp, ventral view
(Kovblyuk et al. 2012b)
Pedipalp, retrolateral view
(Bosmans et al. 2018)
Pedipalp, ventral view
(Bosmans et al. 2018)
Pedipalp, ventral view
(Bosmans et al. 2018)
Pedipalp, ventral view
(Løvbrekke unpubl.)
Pedipalp, ventral view
(Breitling 2020)
Pedipalp, ventral view
(Grimm 1985)
Pedipalp, prolateral view
(Ponomarev & Shmatko 2023b)
Pedipalp, prolateral view
(Ponomarev & Shmatko 2023b)
Pedipalp, retrolateral view
(Ponomarev & Shmatko 2023b)
Tip of pedipalp
(Locket & Millidge 1951)
Bulbus, apical view
(Kovblyuk et al. 2012b)
Bulbus, apical view
(Kovblyuk et al. 2012b)
Bulbus, dorsal view
(Šestáková et al. 2022)
Bulbus, ventral view
(Šestáková et al. 2022)
Bulbus, ventro-prolateral view
(Šestáková et al. 2022)
Bulbus, apical view
(Ponomarev & Shmatko 2023b)
Bulbus, dorsal view
(Ponomarev & Shmatko 2023b)
Bulbus, ventral view
(Ponomarev & Shmatko 2023b)
Embolic apophyis, ventral view
(Bosmans et al. 2018)
Embolic apophyis, ventral view
(Bosmans et al. 2018)
Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)
Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)
Palpal tibia, dorso-lateral view
(Løvbrekke unpubl.)
Palpal tibia, retrolateral view
(Breitling 2020)
Tibial apophysis
(Brændegaard 1966)
Tibial apophysis
(Noordam 1992)
Tibial apophysis
(Locket & Millidge 1951)
Tibial apophysis, retrolateral view
(Kovblyuk et al. 2012b)
Epigyne
(Brændegaard 1966)
Epigyne
(Noordam 1992)
Epigyne
(Roberts 1995)
Epigyne
(Locket & Millidge 1951)
Epigyne, ventral view
(Kovblyuk et al. 2012b)
Epigyne
(Bosmans et al. 2018)
Epigyne
(Bosmans et al. 2018)
Epigyne
(Løvbrekke unpubl.)
Epigyne
(Morka unpubl.)
Epigyne
(Ponomarev & Shmatko 2023b)
Vulva
(Platnick & Shadab 1975)
Vulva, ventral view
(Tullgren 1946)
Vulva, dorsal view
(Kovblyuk et al. 2012b)
Vulva, dorsal view
(Bosmans et al. 2018)
Vulva
(Bosmans et al. 2018)
Vulva
(Løvbrekke unpubl.)
Tibia IV, dorsal view
(Løvbrekke unpubl.)
Tibia I, retrolateral view
(Løvbrekke unpubl.)
Tibia IV, retrolateral view
(Løvbrekke unpubl.)
Tibia I, ventral view
(Løvbrekke unpubl.)
Metatarsus I, ventral view
(Løvbrekke unpubl.)
Distribution List
"No references" does not mean that the species does not occur in this country, but that we have not yet inserted the reference for it. We are working on it.
References
Aakra K, Hauge E (2003) Checklist of Norwegian spiders (Arachnida: Araneae), including Svalbard and Jan Mayen. Norwegian Journal of Entomology 50: 109-129 
Agnarsson I (1996) Íslenskar köngulær Fjölrit Náttúrufræðistofnunar 31: 1-175
Arnold K (2001) Beitrag zur Spinnenfauna (Arachnida, Araneae) des Fürstentums Liechtenstein. Berichte der Botanisch-Zoologischen Gesellschaft Liechtenstein-Sargans-Werdenberg 28: 211-244
Benhacene R, Adjami A, Hadjeb A, Kermiche K, Ouakid M L (2023) Bibliographic checklist of the Algerian spider fauna (Araneae). Zootaxa 5352: 301-357
Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018)
Bosmans R, Kherbouche-Abrous O, Benhalima S, Hervé C (2018) The genus Haplodrassus Chamberlin, 1922 in the Mediterranean and the Maghreb in particular (Araneae: Gnaphosidae) Zootaxa 4451: 1-67
Bosmans R, Van Keer J, Russell-Smith A, Hadjiconstantis M, Komnenov M, Bosselaers J, Huber S, McCowan D, Snazell R, Decae A, Zoumides C, Kielhorn K-H, Oger P (2019c) Spiders of Cyprus (Araneae). A catalogue of all currently known species from Cyprus. Newsletter of the Belgian Arachnological Society 34, Suppl. 1: 1-173
Bosmans R, Van Keer K (2017) Een herziene soortenlijst van de Belgische spinnen (Araneae). Nieuwsbrief van de Belgische Arachnologische Vereniging 32: 39-69
Bosmans R. (unpubl.) Provisional list of spiders of North Africa. Database excerpt Aug. 2019
Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). Zootaxa 4614: 201-254
Breitling R (2020) South European spiders from the Duffey collection in the Manchester Museum (Arachnida: Araneae). Arachnology 18: 333-362
Brændegaard J (1966) Edderkopper: Eller Spindlere I Danmarks Fauna 72: 1-224
Cera I (2018) The checklist of Latvian spiders (Arachnida: Araneae). Environmental and Experimental Biology 16: 139-152
Danışman T, Kunt K B, Özkütük R S, Coşar İ (2024a) The checklist of the spiders of Turkey. Version 2024 [last updated 01 January 2024], online at http://www.spidersofturkey.info
Deltshev C, Vrenozi B, Blagoev G, Lazarov S (2011b) Spiders of Albania – faunistic and zoogeographical review (Arachnida: Araneae). Acta Zoologica Bulgarica 63: 125-144
Demircan Aksan N, Topçu A (2022c) A preliminary checklist of the spider fauna of European Türkiye. Serket 19: 39-56
Febrer J B, Barrientos J A (2022) Ocho primeras citas de arañas (Araneae) para Menorca (Islas Baleares, España). Revista Ibérica de Aracnología 40: 137-144
Grimm U (1985) Die Gnaphosidae Mitteleuropas (Arachnida, Araneae). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg (NF) 26: 1-318
Hermann E (1998) Die Spinnen (Araneae) ausgewählter Halbtrockenrasen im Osten Luxemburgs. Bulletin de la Société des naturalistes Luxembourgeois 99: 189-199
Ivanov V V (2013) The checklist of Belarusian spiders (Arachnida, Araneae). Zoology and Ecology 4: 293-311
Komnenov M (2014) Spider fauna of the Osogovo Mt. Range, Northeastern Macedonia. Fauna Balkana 2: 1-267
Koponen S, Fritzén N R, Pajunen T (2016) Checklist of spiders in Finland (Araneae). 6th version, December 2016, online at http://biolcoll.utu.fi/arach/checklist_of_spiders_in_Finland.htm
Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91
Kovblyuk M M, Kastrygina Z A, Omelko M M (2012b) A review of the spider genus Haplodrassus Chamberlin, 1922 in Crimea (Ukraine) and adjacent areas (Araneae, Gnaphosidae). Zookeys 205: 59-89
Kreuels M, Staudt A, Christian S (2019) Die Spinnenfauna von Luxemburg – eine Zusammenstellung der Nachweise aus den Jahren 1906-2018 (Arachnida: Araneae). Bulletin de la Société des naturalistes luxembourgeois 121: 203-230
Kronestedt T (2001) Checklist of Spiders (Araneae) in Sweden [Preliminary version February 2001]
Lavery A (2019) A revised checklist of the spiders of Great Britain and Ireland. Arachnology 18: 196-212
Le Peru B (2007) Catalogue et répartition des araignées de France. Revue Arachnologique 16: 1-468
Lissner J, Jensen J-K, Hansen L J, Simonsen W, Kelduni R, Nissen K (2016b) An updated checklist of spiders (Araneae) of the Faroe Islands. Norwegian Journal of Entomology 63: 197-240
Locket G H, Millidge A F (1951) British spiders. Vol. I. Ray Society, London, 310 pp.
Løvbrekke H (unpubl.)
Milosevic B (2002) Aranea – Popis vrsta – Checklist. Hrvatsko entomološko društvo – Inventar entomofaune Hrvatske [Aranea – List of species – Checklist. Croatian Entomological Society – Inventory of Croatian entomofauna].
Morka G H (unpubl.)
Naumova M, Lazarov S, Deltshev C (2019a) Faunistic diversity of the spiders in Montenegro (Arachnida: Araneae). Ecologica Montenegrina 22: 50-89
Noordam A P (1992) Gnaphosidae (Araneae) of the Netherlands, drawings, some notes, and some thoughts about ant-mimicry. viii & 16 pp. Published by the author.
Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.
Pantini P, Isaia M (2019) Araneae.it: the online catalog of Italian spiders, with addenda on other arachnid orders occurring in Italy (Arachnida: Araneae, Opiliones, Palpigradi, Pseudoscorpionida, Scorpiones, Solifugae). Fragmenta Entomologica 51: 127-152
Pfliegler W A, Schönhofer A, Niedbała W, Vella P, Sciberras A, Vella A (2017) New records of mites (Acari) and harvestmen (Opiliones) from Malta with a preliminary checklist of Maltese Arachnida. Soil Organisms 89: 85-110
Platnick N I, Shadab M U (1975) A revision of the spider genera Haplodrassus and Orodrassus (Araneae, Gnaphosidae) in North America. American Museum Novitates 2583: 1-40
Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App
Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp.
Ponomarev A V, Shmatko V Y (2023b) Two new species and new records of spiders (Aranei) in Dagestan (Russia). Caucasian Entomological Bulletin 19: 213-219
Roberts M J (1995) Collins Field Guide: Spiders of Britain & Northern Europe. HarperCollins, London, 383 pp.
Rozwałka R, Stanska M (2008) Check-list of spiders (Araneae) of Poland. 1. December 2008. online at http://www.arachnologia.edu.pl/en/species/8-spiders-of-poland.html
Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html
Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184
Šestáková A, Černecká Ľ, Naumova M, Purgat P, Szita É, Gajdoš P (2022) A review of two very rare ground spiders from sandy habitats, new for Slovakia (Araneae: Gnaphosidae). Arachnologische Mitteilungen 64: 14-24
Tullgren A (1946) Svensk spindelfauna. 3. Egentiliga spinddlar. Araneae Fam 5-7. Clubionidae, Zoridae och Gnaphosidae. Stockholm (Entomol Fören.): 141 pp.
Tuneva T K, Esyunin S L (2003a) A review of the Gnaphosidae fauna of the Urals (Aranei), 3. New species and new records, chiefly from the south Urals. Arthropoda Selecta 11: 223-234
Vasiliev A I, Mikhailov K G (2023) Annotated checklist of spiders (Araneae) of the Republic of Moldova. Invertebrate Zoology 20: 307-342
Vilbaste A (1987) Eesti Ämblikud (Aranei). Annoteeritud nimestik. Estonian spiders (Aranei). An annotated check list. Valgus, Tallinn. 113 pp. & 512 maps on 67 unpag. pages
Vrenozi B, Jäger P (2013) Spiders (Araneae) from Albania and Kosovo in the collection of Carl Friedrich Roewer. Arachnologische Mitteilungen 46: 17-26
WSC (2024) World Spider Catalog. Version 25.5. Natural History Museum Bern, online at http://wsc.nmbe.ch (31.10.2024) doi: 10.24436/2
Updates
| 12-01-2024 | Image insert | |
| 05-12-2023 | Image insert | |
| 20-03-2023 | Image insert | |
| 15-01-2023 | Distribution update | Detail |
| 21-11-2022 | Distribution update | Detail |
| 23-10-2022 | Gallery image insert | |
| 26-09-2022 | Image insert | |
| 01-07-2022 | Distribution insert | Detail |
| 19-06-2021 | Distribution update | Detail |
| 29-01-2021 | Distribution insert | Detail |
| 26-08-2020 | Distribution update | Detail |
| 04-05-2020 | Image insert | |
| 17-03-2020 | Distribution insert | Detail |
| 02-10-2018 | Image insert | |
| 10-09-2018 | Image insert |